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Public fears are high that foreign agents are among us, waiting for the order to strike. A divided America is debating whether or not to go to war. The FBI and police, scrambling to thwart any attacks, round up the plotters. Experts declare that our intelligence capabilities are insufficient and that a new agency must be created.

The year is 1941.

Fifth Column is the story of Johanna Falck, a German immigrant who joins the new American central intelligence service. As Americans focus on the war in Europe and whether the United States should intervene, the FBI is rounding up scores of German spies. The German-American Bund, a pro-Nazi group of Americans suspected of being saboteurs and subversives, is at the center of the FBI's investigations.

Johanna is recruited to infiltrate the Bund and discover what they and the Nazis have planned. Soon she is caught up in a far-reaching conspiracy, one that stretches from the top of the Nazi state to the streets of New York.

What she finds shatters her most basic assumption about the Third Reich.